Tag: prosauropod

It Came From The Trash Heap (We Don’t Talk About Kholumolumo)

A wastebasket taxon is what happens when species can’t be easily classified and instead get hurled into a “catch-all” category.

…But that’s not the only kind of taxonomic tangle that can befall a new discovery.

When a scientific name is assigned to a new species, but it isn’t given a corresponding formal description and type specimen, it becomes a nomen nudum – a “naked name”. Without a proper description and assigned holotype the name isn’t valid, and the new species isn’t technically accepted by the wider scientific community.

This has even happened to some surprisingly famous names. In the 1920s Velociraptor mongoliensis was briefly given the nomen nudum “Ovoraptor djadochtari” before getting its much more familiar name when it was officially described. Meanwhile the giant pterosaur Quetzalcoatlus northropi was stuck as a nomen nudum for decades, only finally getting a proper published description in 2021.

And there’s another particular long-standing nomen nudum that became mildly infamous – “Thotobolosaurus”, the “trash heap lizard”.

An illustration of Kholumolumo (previously known as "Thotobolosaurus"), an extinct prosauropod dinosaur. It's a bipedal dinosaur with a small head, long neck, chunky arms, thick bird-like legs, and a long counterbalacing tail. It's colored dark blue, with lighter striping on its underside and a speckled red-and-orange pattern along its back. Its face is brighter teal-blue with a mask-like streak of orange than continues down onto a fleshy throat dewlap. There are also reddish quill-like filaments on the back of its neck and in a tuft at the tip of its tail.
Kholumolumo ellenbergerorum

Discovered next to a literal trash pile in the village of Maphutseng in Lesotho, a few scattered and broken bones of this “prosauropod” sauropodomorph dinosaur were first found in 1930. But it wasn’t until the mid-1950s that a more extensive bonebed began to be unearthed at the site, and over the next decade over 1000 fossil fragments were collected.

In the mid-1960s the remains were initially classified as belonging to Euskelosaurus browni (which is now considered to be a wastebasket taxon), but just a few years later in 1970 the “Maphutseng Beast” was re-evaluated as a species new to science. It was referred to as “Thotobolosaurus mabeatae” – based on the local name of the discovery site, “Thotobolo ea ‘Ma-Beata” (trash heap of Beata’s mother) – but this name was never actually formally published.

Despite “Thotobolosaurus” being an undescribed nomen nudum it nonetheless went on to be repeatedly referenced in scientific literature over the next few decades, and appeared in several popular dinosaur books (even as recently as 2020!).

In the mid-1990s it was alternatively named “Kholumolumosaurus ellenbergerorum” in a Ph.D. dissertation, with this name derived from the kholumolumo, a reptilian creature in Sotho mythology, and the Ellenberger brothers who worked on the site. But this also didn’t count as a formal publication and instead became a second nomen nudum for the species.

Eventually, 90 years after the first bones were found and 50 years after the debut of the name “Thotobolosaurus”, this long-neglected sauropodomorph was finally given a proper published full anatomical description in 2020.

And it also got a third name, this time officially valid, based on the second one from the 1990s: Kholumolumo ellenbergerorum.

For something associated with trash for so long, Kholumolumo is actually now one of the most completely-known prosauropods. At least five different individuals were present in the collected fossil material, possibly as many as ten, and between them most of the full skeleton is represented – with the exception of the skulls, which are only known from a couple of small fragments.

We now know Kholumolumo was rather heavily-built, with chunky limb bones and unusually short shinbones. It would have been one of the biggest animals around in the Late Triassic (~210 million years ago), measuring at least 9m long (~30′) and weighing around 1.7 tonnes (1.9 US tons), but despite its size it seems to have still been bipedal.

Due to the highly disarticulated nature of the bones the fossil site may have been a “bone accumulation area”, a place where dismembered bits and pieces of different carcasses were regularly carried to be eaten by a predator or scavenger  – essentially a trash heap, fittingly enough. A couple of “rauisuchian” teeth have actually been found among the remains, which might indicate what was chomping on these particular Kholumolumo.

Retro vs Modern #10: Plateosaurus trossingensis

First discovered in southeast Germany in the 1930s, Plateosaurus was only the fifth non-avian dinosaur known to science – but its fossils were fragmentary and poorly understood until the early 20th century, when large bonebeds full of much better specimens began to be excavated.

1910s-1970s

Between the 1910s and 1930s around 80 near-complete skeletons of Plateosaurus were found in two German quarries, quickly making it one of the most abundant and best known dinosaur species of the time. Although it had previously been classified as a theropod dinosaur, in the 1920s the more complete material allowed it to be properly identified as a “prosauropod“, an early herbivorous relative of the giant sauropods. 

Like many bipedal dinosaurs during this period Plateosaurus was generally interpreted as having an upright kangaroo-like posture with a dragging tail – although some paleontologists were arguing for it having a sprawling quadrupedal lizard-like stance as late as the mid-1930s.

1980s

Unfortunately much of the German fossil material was destroyed during World War II bombing raids, and interest in Plateosaurus didn’t pick up again until the time of the Dinosaur Renaissance when a third major fossil site was discovered in Switzerland during the 1970s.

Plateosaurus was reinterpreted with a horizontal body posture and fully upright limbs. As an early member of the sauropodomorph lineage it was often depicted as a transitional form between bipedal ancestral dinosaurs and the later quadrupedal sauropods, thought to primarily walk on all fours but also able to run on its hind legs like a hadrosaur – although some studies instead concluded it was fully quadrupedal with a downwards-curling tail that made bipedal movement impossible.

The large numbers of skeletons found together were considered to represent evidence for herding behavior, with groups of Plateosaurus being caught in catastrophic mudflows all at once.

2020s

Extensive biomechanical studies in the 2000s and early 2010s clarified what sort of posture Plateosaurus was really capable of. It was found to be completely unable to position its arms in a quadrupedal stance, and so was actually purely bipedal – and skeletons that had been mounted in the quadrupedal position had needed many of their joints to be completely dislocated to achieve the pose!

A huge number of different Plateosaurus species had been named over the genus’ nearly-200-years of history, too, creating a confusing mess of dubious and invalid names. These were all finally revised in 2019 leaving just three valid species, with Plateosaurus trossingensis as the best known and the new type species.

We now know Plateosaurus lived across central and northern Europe during the Late Triassic, around 214-204 million years ago, at a time when the region had a subtropical climate. It had a small head on a long flexible neck, with teeth convergently resembling those of modern iguanas suggesting it was probably primarily herbivorous (with possible opportunistic omnivory). Its arms were proportionally short for a prosauropod but were well-adapted for grasping, with large claws that may have been used to dig up roots and tear down branches

It had a rapid growth rate and bird-like lungs and air sacs that suggest it was warm-blooded, and different individuals showed an unusually high amount of variation in adult size and age of maturity. Some appear to have been fully grown at about 5m long (~16′) and as young as 12 years old, while others reached 10m long (~33′), and were still growing at 27 years old.

The bonebeds are no longer thought to represent mass mortalities of herds, but instead were probably a scenario more similar to the La Brea Tar Pits – mud-miring traps that smaller lighter animals could escape from but larger individuals became stuck and died.

No prosauropod skin impressions have been found yet, so it’s still unknown whether Plateosaurus was scaly like later sauropods or if it had some degree of protofeather hair-like fuzz.

Island Weirdness #01

Islands are natural sites for evolutionary experiments. Their isolation and limited resources put a lot of selective pressure on their native species, often resulting in spectacular and unique adaptations. Big animals become small, small animals become big, and ecological niches can end up being filled in unexpected ways.

From the dodo becoming the first well-known example of human-caused extinction, to Darwin’s Galápagos finches being influential in the development of the theory of natural selection, to famous-but-endangered living examples like the kiwi and marine iguana, island species are fascinating and often fragile examples of how diverse life can get even in restricted conditions.

In fact, this theme ended up containing so many species I wanted to feature that I can’t possibly fit them all into just a single month. So, for the first time, a theme is going to need two months – with part 1 happening right now, and part 2 coming later this summer.

Thecodontosaurus antiquus

For much of the Mesozoic Europe was an archipelago of islands in a shallow tropical sea. During the Late Triassic, about 205-201 million years ago, some of the paleo-islands in this region existed around southern Wales and South West England, near the city of Bristol.

Thecodontosaurus was actually one of the first non-avian dinosaurs ever named by modern science, discovered in the mid 1830s – several years before the term “dinosaur” was even created to classify the “great ancient lizards”.

It was an early member of the herbivorous sauropodomorphs, the group that would eventually include the largest ever land animals. But unlike its enormous later cousins it was short-necked and bipedal, and was particularly small compared to other contemporary “prosauropods”, measuring only about 2m long (6′6″). This would make it one of the oldest known examples of insular dwarfism.