Cambrian Explosion #53: Trilobita – A Prolific Paleozoic Posse

The biggest stars of the Cambrian euarthropods, and most of the Paleozoic Era, were of course the trilobites. Known from literally tens of thousands of species spanning over 270 million years, they’re some of the most recognizable and popular fossils.

Trilobites’ exact evolutionary origins and transitional forms are unknown, but they’re thought to have originated in Siberia in the very early Cambrian and their leg anatomy indicates they were a part of the artiopodan lineage. They made a sudden and dramatic entrance to the fossil record about 521 million years ago, appearing fully-formed and rapidly diversifying and spreading all around the world within just a couple of million years.

Their hard calcified exoskeletons made them much more likely to fossilize than soft-bodied animals, with a distinctive three-part body plan consisting of a head shield, three-lobed thorax segments, and a tail shield. Each individual regularly molted their carapace throughout their life, meaning that most trilobite remains are actually empty discarded shells rather than actual carcasses.

Along with being heavily armored arthropod tanks, most species were also able to roll themselves up to defend against predators, and some developed additional elaborate spines and spikes.

…And some were just weird.

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