amphibian – Page 3 – Nix Illustration

The newly-named Chinlestegophis jenkinsi, in the style of last year’s Amphibian August illustrations.

Living during the Late Triassic of Colorado, USA, (~220 mya), this 30cm long (1′) amphibian had a skull showing a mixture of features shared with both temnospondyls and modern caecilians – providing a vital “missing link” in their evolutionary history. Previously the oldest known caecilian-relative was the Jurassic-aged Eocaecilia, which already had much more modified anatomy making it harder to definitively link to other known groups.

Chinlestegophis seems to have been part of the stereospondyl branch of the temnospondyls – and an unexpected side effect of adding caecilians into this group is that many temnospondyls could now potentially also be classified as true members of Lissamphibia.

An example family tree showing this new version of amphibian relationships. If we define Lissamphibia as including all living amphibians and their extinct relatives back to their last shared common ancestor, then everything within the grey box is a lissamphibian!

Of course, this is still just one hypothesis of amphibian evolution among several other competing ideas. Maybe it’s right, maybe it isn’t – as always, we need more fossil evidence! – but it’s certainly an interesting and surprising new development in the ongoing saga of “what are lissamphibians? we just don’t know”.

2023 Update: …But the discovery of Funcusvermis suggests Chinlestegophis isn’t a caecilian at all, but instead an unrelated case of convergent evolution!

Sirens are part of the salamander branch of the lissamphibians, and are some of the most unusual members of the group. They have an eel-like body shape, with small forelimbs and no hindlimbs at all, and have functional external gills as adults. Their main diet is carnivorous, with palatal teeth and a keratinous beak at the front of their jaws adapted for eating hard-shelled prey – but they’ve also been observed feeding on plant material, a rarity among amphibians.

Habrosaurus here was one of the earliest known sirens, living during the Late Cretaceous and Early Paleocene of North America (~84-58 mya). Reaching lengths of around 1.6m (5′3″), it was one of the largest known lissamphibians of all time, comparable in size to some modern giant salamanders.

It lacked the beak seen in its modern relatives, instead having specialized chisel-like teeth at the edges of its jaws that convergently served the same purpose of delivering a crushing bite.