Turtle-jawed moa-nalo

The turtle-jawed moa-nalo (Chelychelynechen quassus) was a large flightless goose-like duck from the Hawaiian island of Kaua‘i. About 90cm tall (3′) and weighing around 7kg (15lbs), these birds and their relatives were descended from dabbling ducks and existed on most of the larger Hawaiian islands for the last 3 million years or so – before going extinct around 1000 years ago following the arrival of Polynesian settlers.

Chelychelynechen had an unusually-shaped bill, tall and broad with vertically-oriented nostrils, convergently similar to the beak of a turtle. It would have occupied the same sort of ecological niche as giant tortoises on other islands, filling the role of large herbivore in the absence of mammals.

Bathornis

Bathornis grallator, a flightless bird about 75cm tall (2′6″) from the Late Eocene and Early Oligocene of Midwestern USA (~37-34 mya).

It was originally mistaken for a long-legged vulture (under the name Neocathartes) when first discovered in the 1940s, but later studies have shown it was actually one of the smaller members of the bathornithids – close cousins of the more well-known South American “terror birds”, successfully occupying terrestrial predator niches alongside large carnivorous mammals.